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The relations between organisms and places where they live, together with the atmosphere, can be classified in different Ecological niches. In the underground environment we can identify many different kinds of habitats.

Ecological Classification of the Environments


Cave environment. This is a place accessible to man, and can be easily visited without eany problem.
Interstitial environment. It is a labirinthic part of the underground with many microfissures wich man can not visit.
Endogean environment. Region between the inferior limit of the vegital detritus and the inferior limit of the roots of arboreal plants. 
Mammal dens and Formicaries. Interesting habitat for most specialized species.
Phreatic environment. This zone is full of water and is subject to seasonal variations of level, depending on meteriological conditions.
Artificial galleries. These are galleries excavated by man.

The figure below shows a schematic description the Hypogean world with all the habitats.


As showed in the figure above, life has many habitats in the Hypogean world. Animals are most specialized in there environment and for them it is not possible to live in other places. This means that the ecosystem is much influenced by external metereological conditions. Infact rain goes underground and makes it possible for water to reach all kinds of niches. In caves there are many spaces where species can live: in small and big pools, in the ground under stones or in mud, on the caverns wall, inside guano (dung of Bats) etc..All this is classified in the Cave environment. If we penetrate the microfissures in the cave we go in the Interstitial environment, wich is a very interesting and labirintic place where man can not go. It is beleived that many species found in caves truly live in these cracks, from wich irregularly they reach the cave. Many specialized species can also be found in deep buried stones, that are certainly in comunication with interstitial habitats. Sometimes we find the same conditions in artificial galleries, that have the same climatical parameters.
Another zone is the Phreatic environment, this zone is full of water and is subject to seasonal variations of level, depending on meteriological conditions. The connection between the epigean and hypogean system is quit remarkable, due to the influence of rain.
Under the ground, between the inferior limit of the vegital detritus and the inferior limit of the roots of arboreal plants, we distinguish the Endogean environment. Here we find many species and it can be compared with the interstital and caves becouse of high humidity, no light, same temperature etc.. The aspect of this portion of ground can be full of rocks, gravelly, argillaceous and friable  depending on the soil. In high altitudes the thickness can be of some decimeters, alpine pastures, or can reach some meters like in boschive areas.
Also interesting to consider is the life in the dens of mammals, formicaries and termitaries, that sometimes reach a considerable depth under the soil. Also in this case the specialization of species is at high level.

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